Runway 34 tells the turbulent tale of Captain Vikrant Khanna (Ajay Devgn), a pilot prodigy whose flight takes a stirring and mysterious trajectory after take-off from an international destination. Consequently, it follows the tenacious Narayan Vedant (Amitabh Bachchan) in his pursuit to uncover the truth.
The film begins when Gopika and her father move into a house where Bharathiyamma stays with her two children, Balu Damodar and Ambili. Bharathiyamma has one more elder son Unni Damodar who works in Dubai. Unni tells his family that he is working in Gulf but in reality he runs illegal spirit business with Chandy Bhai along with Achayan and Achuvettan in Valayar.
He accepts a four-year imprisonment but actually the accused was Bhai. Unni, however confesses what he does, to Gopika whom his mother is insistent that he should marry her. After leaving the jail Paramasivam realizes that Bhai has fallen into a huge debt because of the cheating of Chinnadan Varkey and his sons. He then goes to Valayar and thus saves Bhai and his family from debt with his wicked mind and tricks. He again rules spirit deals in Valayar area pulling the Chinnadans into trouble. Things change when Balu becomes a police officer. He is posted in Valayar area, where Unni and Bhai have become rowdies.
In Valayar, Balu tries his level best to catch Valayar Paramasivam but does not know that it is none other than his brother; and fails in mind game played by Unni. This makes Bhai angry with Balu. But Paramasivam tells him to forgive Balu because he was his brother. In the meantime, Unni's family discovers that he himself is Paramsivam after Gopika's uncle Divakaran, a jail guard in the jail where he was imprisoned visits them and he is ousted from his family. Later, Chinnadan kills Bhai's only son Johnny and blames Balu as the real killer. This made Bhai very desperate and wants to kill Balu. Bhai kidnaps him. On knowing this, Paramasivam goes to save Balu and informs Bhai that the actual killer was SP by identifying the ring marks in Johnny's face.
Paramasivam then confesses him that Bhai's Paramasivam will not lie to Bhai. After realizing the innocence of Balu, Bhai and Paramasivam plans to kill the Chinnadans by releasing gas and spirit into their house after locking all the doors and sending a toy plane which blasts when functioned switch is pressed. The film moves to the climax, where it is shown that Paramasivam gets acquitted from all their cases since Bhai has pleaded guilty to all the crimes. He wants Paramasivam to return as Unni to his family and lead a normal life with them.
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